The authors would like to thank Bolu Abant Izzet Baysal University for the financial support of the first author to carry out the field studies (project no: BAP-2019.03.01.1428), Dr. Funda Ozbek for SEM photographs, Assoc. Prof. Dr. Mehtap Teksen for her contribution to the interpretation of seed micromorphology and Fatma Sen for drawing the illustration. We are also grateful to the directors and curators of AEF, AIBU, ANK, DUOF, E, G, GAZI, HUB, K, NGBB, Herbarium of Sakarya University, and BM, JE, L, MNHN-P, WU digital herbaria for allowing the first author to examine the Dherbarium samples.
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